schultz

Dr. Tom D. Schultz

AffiliationFaculty
TitleTight Family Professor in the Natural Sciences, Behavioral Ecologist
Office227 Talbot Hall
Email
Phone
740-587-6218
Fax740-587-5634
Ph.D. in Zoology from University of Texas at Austin, 1983
A.B. in Biological Sciences from University of Chicago, 1977
HS diploma from University of Chicago, 1974

Memberships

  • Animal Behavior Society
  • Coleopterist's Society
  • Dragonfly Society of America
  • Ecological Society of America
  • International Society of Behavioral Ecology
  • Ohio Odonata Society
  • Society for the Study of Evolution

Academic Positions

Professor of Biology, Department of Biology at Denison University, 2000 - present
Chair, Department of Biology at Denison University, 1998-2001
Director, Denison University Biological Reserve, 1994-2003
Visiting Lecturer, Department of Biology at Yale University, 1988-1990
Post-doctoral Research Associate, Department of Zoology at Arizona State University, 1985-1987
Lecturer, Department of Zoology at The University of Texas at Austin, 1983-1984

Teaching

I find it very satisfying and fun to explore the natural world, but my real passion is for the learning process. The challenge of being exposed to a new idea, questioning it, evaluating it, and even testing it, is very fulfilling for me. I especially enjoy making connections between seemingly disparate ideas or concepts in different disciplines (an important ability in a time when boundaries between scientific disciplines are becoming blurred). The courses I teach all involve integrating different approaches and levels of organization. Sometimes I think I have the perfect job in that I am paid to learn new things and share them with students, and to help them to develop a "Swiss Army Knife" of critical thinking skills. With these skills, they can become good leaders and thoughtful citizens in any field, and better able to face the uncertainty of the future.

Courses Taught:

First Year Studies: Animal Talk
Introduction to the Science of Biology
Ecology & Evolution
Biology of Insects
Animal Behavior
Senior Research

Research

Interests

  • Sensory Drive and the Evolution of Insect Visual Signals
  • Sexual Selection on Damselfly Coloration
  • The Evolution of Anti-Predator Tactics in Insects
  • Structural Colors and Interference Reflectors in Insects
  • Conservation and Biomonitoring of Insects

I am interested in the evolution of animal coloration in response to visual predators, sexual selection, thermal constraints, and the light environment. Much of my previous work has focused on the influence of habitat preference, foraging behavior, and thermoregulation on the evolution of cryptic defenses in tiger beetles and other insects. Currently, my research is focused on color polymorphisms in damselflies and their effectiveness as intraspecific signals. I am also interested in the evolution of structural colors and iridescence in insects. Finally, I am frequently engaged in biodiversity assessment and monitoring projects using insects as indicator taxa.

Click on images to enlarge.

clepida2.jpg
How do structural colors provide camoflage in tiger beetles?
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Does color influence mate choice in damselflies?
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Are conspicuous colors of male damselflies signals of sexual and species identity?
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How do structural colors provide flashing UV signals in tropical helicopter damselflies?

Publications

Schultz, T.D., C.N. Anderson*, & L. B. Symes*. 2008. The conspicuousness of colour cues in male pond damselflies depends on ambient light and visual system. Animal Behaviour (in press) *.
* Student co-authors
Fincke, O.M., A. Fargevielle, & T. D. Schultz. 2007. Lack of innate preference for morph and species identity in mate-searching Enallagma damselflies. Behavioral Ecology and Sociobiology. v. 61 p. 1121-1131
Fincke, O.M., R. Jodicke, D. Paulson, & T. D. Schultz. 2005. The frequency of female-specific color polymorphisms in Holarctic Odonata: why are male-like females typically the minority? . International Journal of Odonatology. v. 8 p. 183-212
Schultz, T. D. 2001. Tiger beetle defenses revisited: alternative defense strategies and colorations in two neotropical tiger beetles, Odontocheila nicaraguensis and Pseudoxycheila tarsalis (Carabidae: Cicindelinae). Coleopterists Bulletin. v. 55 p. 153-163
Schultz, T. D. & J. Puchalski *. 2001. Chemical Defenses in the Tiger Beetle Pseudoxycheila tarsalis Bates (Carabidae: Cicindelinae). Coleopterists Bulletin. v. 55 p. 164-166
Kirkton, S. D.* & T. D. Schultz. 2001. Age-specific behavior and habitat selection of adult male damselflies, Calopteryx maculata (Odonata: Calopterygidae). Journal of Insect Behavior . v. 14 no. 4 p. 545-556
Schultz, T. D. 1998. The utilization of patchy thermal microhabitats by the ectothermic insect predator, Cicindela sexguttata. Ecological Entomology . v. 23 p. 444-450
Knisley, C. B. & T. D. Schultz. 1997. The Biology of Tiger Beetles and a Guide to the Species of the South Atlantic States. Virginia Museum of Natural History, Martinsville, VA. p. 236 pp
Schultz, T. D., M. Quinlan & N. F. Hadley. 1992. Preferred body temperature, metabolic physiology, and water balance of adult Cicindela longilabris: a comparison of populations from boreal habitats and climatic refugia. Physiological Zoology . v. 65 p. 226-242
Hadley, N. F., A. Savill, & T. D. Schultz. 1992. Coloration and its thermal consequences in the New Zealand tiger beetle Neocicindela perhispida. J. Thermal Biology . v. 17 p. 55-61
Schultz, T. D. 1991. Tiger Hunt. Natural History . p. 38-44
Schultz, T. D.& G. Bernard. 1990. Pointillistic mixing of interference colors in cryptic tiger beetles. Nature . v. 337 p. 72-73
Hadley, N. F., T. D. Schultz, & A. C. Savill. 1988. Spectral reflectances of three subspecies of the tiger beetle Neocicindela perhispida: correlations with their respective habitat substrates. New Zealand J. of Zoology. v. 15 p. 343-346
Schultz, T. D. & N. F. Hadley. 1987. Microhabitat segregation and physiological differences in co-occuring tiger beetle species, Cicindela oregona and Cicindela tranquebarica. Oecologia . v. 73 p. 363-370
Schultz, T. D. & N. F. Hadley. 1987. Structural colors of tiger beetles and their role in heat transfer through the integument. Physiological Zoology . v. 60 p. 737-745
Schultz, T. D. 1986. The role of structural colors in predator avoidance by tiger beetles of the genus Cicindela. Bulletin of the Entomological Society of America . v. 32 p. 142-146
Schultz, T. D. & M. A. Rankin. 1985. The ultrastructure of epicuticular interference reflectors of tiger beetles (Cicindela). J. Experimental Biology . v. 117 p. 88-110
Schultz, T. D. & M. A. Rankin. 1985. Developmental changes in the interference reflectors and colorations of tiger beetles (Cicindela). J. Experimental Biology. v. 117 p. 111-118

Senior Research

One of the best aspects of being at a small college is the opportunity for close faculty-student collaboration on independent research. I have had the pleasure of advising a number of undergraduate students who have conducted a variety of outstanding research projects, many of which have been presented at national scientific meetings. As a research advisor, I involve students in my studies of insect behavioral ecology or enlist students interested in conservation biology in conducting inventories and monitoring studies at the Bio Reserve and other sites in Licking County.  In almost all cases, these projects require a summer of field work prior to the senior year. In the past, my summer research students have been supported with Anderson Research Fellowships or stipends provided through the Denison University Research Foundation.

Research Projects Supervised

Brindle, A. 2008. Differing social environments between primate populations may generate false positive evidence of cultural variation. *

Gorsich, E. 2008. Ommochrome signaling in male Enallagma damselflies: can long wavelength coloration be correlated with territorial behavior? *

Bring, B. 2007. A study of odonate community development, habitat preferences, and colonization among ponds and artificial wetlands at Dawes Arboretum.

Horn, J. 2007. Suburban habitat fragmentation: effects on migratory and residential songbirds in Central Ohio. *

Symes, L. 2007. Polychromatism and sex identity signals in the damselfly genus Enallagma. *

Symes, L. 2006. Polychromatism in the damselfly Enallagma civile and an assessment of the Male-Mimicry Hypothesis. (Poster presented at 2006 Meeting of the Ecological Society of America).

Hughes, D. 2005. The interspecific roosting behaviors of the Turkey Vulture (Cathartes aura) and Black Vulture (Corapgys atratus).

Dunlevy, J. 2003. Investigations of summer bird residents at five sites within Licking County.

Anderson, C. 2002. Enallagma damselfly colors as visual signals in relation to ambient light and visual backgrounds. * (Poster presented at 2002 Meeting of the Ecological Society of America).

Bucci, L. 2002. Effects of vegetation and landscape on butterfly diversity and abundance. *

Clark, E. 2002. Corellations of odonate diversity with lotic habitat characteristics.

Menninger, H. 2000. Examining the ecology of an indicator taxon: damselfly species diversity and the role of habitat heterogeneity. * (Paper presented at 2000 Meeting of Ohio Odonata Society).

Hauck, A. 1999. Correlation between male damselfly colorations and the light environments of courtship arenas. *

Menninger, H. 1999. Initiation of a long-term monitoring program for Odonata at the Denison University Biological Reserve. (Poster presented at 1999 Meeting of the Entomological Society of America).

Kirkton, S. 1997. Babes in the woods: age-specific dispersal in the territorial damselfly, Calopteryx maculata. * (Paper presented at 1997 Meeting of the Ecological Society of America).

Scheub, C. 1997. Diversity and abundance of the Papilionoidea at the Denison University Biological Reserve.* (Poster presented at 1996 Meeting of the Entomological Society of America).

Kirkton, S. 1996. Why do male ebony jeweling damselflies (Calopteryx maculata) aggregate far from territorial breeding sites? (Poster presented at 1996 Meeting of the Entomological Society of America).

Casey, J. 1996. The significance of interference colors and visual communication in Phidippus audax, the daring jumping spider. *

Godfrey, P. 1996. Optimal site choice and foraging posture of the ambush predator Phymata fasciatus. *

Forbes, B. 1995. Substrate matching and cryptic defenses in the toad bug Gelastocoris oculatus. *

Stocker, E. 1995. Spectral sensitivity of the visual system in the praying mantis, Sphrodomantis lineola. (Paper presented at the 1994 Meeting of the Entomological Society of America).

Angalich, L. 1994. Correlation between conspicuousness and escape flight behavior among species of tiger beetles. * (Poster presented at the 1993 Meeting of the Entomological Society of America).

Gallo, T. 1994. A review of the history and efficacy of the Endangered Species Act and prospect for its renewal. *

Price, C. 1994. A comparative study of insect colonization and decomposition of pig carrion in central Ohio. * (Paper presented at the 1994 Meeting of the Ohio Academy of Science).

Puchalski, J. 1994. Comparative chemical analysis of the defensive secretions of tiger beetles (Cicindelidae) and leave beetles (Chrysomelidae). *

Van Antwerp, A. 1994. Light gap utilization and behavioral thermoregulation by the green forest tiger beetle, Cicindela sexguttata. * (Poster presented at the 1993 Meeting of the Entomological Society of America).

* denotes Honors Project

Memberships

  • Animal Behavior Society
  • Coleopterist's Society
  • Dragonfly Society of America
  • Ecological Society of America
  • International Society of Behavioral Ecology
  • Ohio Odonata Society
  • Society for the Study of Evolution